Research Interests: Rous sarcoma virus (RSV) and other avian sarcoma and leukosis viruses (ASLV), cell transformation by RSV, v-src oncogene, transcriptinal control of integrated proviruses, DNA methylation, antivirus immune response and immunogenetics of chicken, ASLV-induced pathogenesis in chicken and duck, human endogenous retroviruses and retroelements, cell receptors for ASLV, site-specific integration of retroviruses, carbonic anhydrase IX.